PMID-sentid Pub_year Sent_text comp_official_name comp_offsetprotein_name organism prot_offset 15807788-2 2005 In vertebrates, these functions are controlled by a three-dimensional gradient of Ran-GTP to Ran-GDP, established by the spatial separation of Ran GTPase-activating protein (RanGAP) and the Ran guanine nucleotide exchange factor RCC1. Guanosine Diphosphate 97-100 regulator of chromosome condensation 1 Homo sapiens 229-233